Caio Brunharo, Ph.D.

Caio Brunharo, Ph.D.

  • Assistant Professor of Weed Science
116 Agriculture Science and Industries Building
University Park, PA 16802


  • BS Agricultural Engineering (2013) University of Sao Paulo
  • MSc Crop Science (2014) University of Sao Paulo
  • PhD Horticulture and Agronomy (2018) University of California, Davis
  • Google Scholar:


Modeling weed seedling emergence for time-specific weed management: A systematic review
Weed Science, Marschner, Caroline A., Colucci, Isabella, Stup, Rebecca S., Westbrook, Anna S., Brunharo, Caio A.C.G., Ditommaso, Antonio, Mesgaran, Mohsen B., 2024

Evolution of ACCase-inhibitor resistance in Chloris virgata is conferred by a Trp2027Cys mutation in the herbicide target site
Pest Management Science, Ferreira, Luiz Augusto Inojosa, de Oliveira, Rubem Silvério, Constantin, Jamil, Brunharo, Caio, 2023

First report of target-site resistance to ACCase-inhibiting herbicides in Bromus tectorum L.
Pest Management Science, Ribeiro, Victor Hugo Vidal, Brunharo, Caio A.C.G., Mallory-Smith, Carol, Walenta, Darrin L., Barroso, Judit, 2023

Herbicide resistance in Bromus spp.: a global review
Weed Science, Ribeiro, Victor H.V., Barroso, Judit, Brunharo, Caio A.C.G., Mallory-Smith, Carol, 2023

Western United States and Canada perspective: are herbicide-resistant crops the solution to herbicide-resistant weeds?
Weed Science, Brunharo, Caio A.C.G., Gast, Roger, Kumar, Vipan, Mallory-Smith, Carol A., Tidemann, Breanne D., Beckie, Hugh J., 2022

Multiple evolutionary origins of glyphosate resistance in Lolium multiflorum
Evolutionary Applications, Brunharo, Caio A.C.G., Streisfeld, Matthew A., 2022

Seed Production, Dissemination, and Weed Seedbanks, Netto, Acacio Goncalves, Christoffoleti, Pedro, VanGessel, Mark, Carvalho, Saul J.P., Nicolai, Marcelo, Brunharo, Caio, 2022

Population genomics of Digitaria insularis from soybean areas in Brazil
Pest Management Science, Gonçalves Netto, Acácio, Cordeiro, Erick M.G., Nicolai, Marcelo, de Carvalho, Saul J.P., Ovejero, Ramiro Fernando Lopez, Brunharo, Caio A.C.G., Zucchi, Maria I., Christoffoleti, Pedro J., 2021

Review: evolutionary drivers of agricultural adaptation in Lolium spp.
Pest Management Science, Matzrafi, Maor, Preston, Christopher, Brunharo, Caio Augusto, 2021

Non-target-Site Resistance in Lolium spp. Globally: A Review
Frontiers in Plant Science, Suzukawa, Andréia K., Bobadilla, Lucas K., Mallory-Smith, Carol, Brunharo, Caio A.C.G., 2021

Season-long weed control with sequential herbicide programs in California tree nut crops
Weed Technology, Brunharo, Caio A.C.G., Watkins, Seth, Hanson, Bradley D., 2020

Mechanism of clomazone resistance in Leptochloa fusca spp. fasicularis to clomazone
Pesticide Biochemistry and Physiology, Driver, Katie E., Brunharo, Caio A.C.G., Al-Khatib, Kassim, 2020

Increased temperatures and elevated CO 2 levels reduce the sensitivity of Conyza canadensis and Chenopodium album to glyphosate
Scientific Reports, Matzrafi, Maor, Brunharo, Caio, Tehranchian, Parsa, Hanson, Bradley D., Jasieniuk, Marie, 2019

Role of Glutamine Synthetase Isogenes and Herbicide Metabolism in the Mechanism of Resistance to Glufosinate in Lolium perenne L. spp. multiflorum Biotypes from Oregon
Journal of Agricultural and Food Chemistry, Brunharo, Caio A.C.G., Takano, Hudson K., Mallory-Smith, Carol A., Dayan, Franck E., Hanson, Bradley D., 2019

EPSPS duplication and mutation involved in glyphosate resistance in the allotetraploid weed species Poa annua L.
Pest Management Science, Brunharo, Caio Augusto De Castro Grossi, Morran, Sarah, Martin, Katie, Moretti, Marcelo L., Hanson, Bradley D., 2019

Multiple target site resistance to glyphosate in junglerice (Echinochloa colona) lines from California orchards
Pest Management Science, Morran, Sarah, Moretti, Marcelo L., Brunharo, Caio A., Fischer, Albert J., Hanson, Bradley D., 2018

Sourgrass resistance mechanism to the herbicide glyphosate
Advances in Weed Science, Melo, Marcelo, Rocha, Lucas, Brunharo, Caio, Nicolai, Marcelo, Tornisiello, Valdemar, Nissen, Scott, Christoffoleti, Pedro, 2018

Resistência de plantas daninhas a herbicidas inibidores da EPSPs (Grupo G), Christoffoleti, Pedro, Nicolai, Marcelo, Melo, Marcel, Silva, Danielo, Brunharo, Caio, 2018

Phosphorus fertilization in sugarcane cultivation under different soil managements
Revista Brasileira de Engenharia Agricola e Ambiental, de Sousa Junior, Paulo R., Brunharo, Caio A.C.G., Furlani, Carlos E.A., Prado, Renato de M., Maldonado Júnior, Walter, Zerbato, Cristiano, 2017

Vacuolar sequestration of paraquat is involved in the resistance mechanism in lolium perenne L. Spp. multiflorum
Frontiers in Plant Science, Brunharo, Caio A.C.G., Hanson, Bradley D., 2017

Chemical control alternatives for glyphosate-resistant sourgrass using hebicides indicated for corn and cotton crop
Brazilian Journal of Herbicides, Melo, Marcel, Rocha, Leonardo, Brunharo, Caio, Silva, Danilo, Nicolai, Marcelo, Christoffoleti, Pedro, 2017

Confirmation and mechanism of glyphosate resistance in tall windmill grass (Chloris elata) from Brazil
Pest Management Science, Brunharo, Caio Acg, Patterson, Eric, Carrijo, Daniela, de Melo, Marcel, Nicolai, Marcelo, Gaines, Todd, Nissen, Scott, Christoffoleti, Pedro, 2016

Manejo de plantas daninhas do gênero Amaranthus na cultura da cana-de-açúcar, Brunharo, Caio, Christoffoleti, Pedro, Christoffoleti, Pedro, 2015

Physiological characterization of sugarcane varieties under oxidative stress caused by the herbicide paraquat
Cientifica, Pavani, Alan, Guimaraes, Ana, Brunharo, Caio, Christoffoleti, Pedro, 2015

Manejo de plantas daninhas, Christoffoleti, Pedro, Brunharo, Caio, Melo, Marcel, Nicolai, Marcelo, Romano, Mylena, 2014

Aspects of the mechanism of action of the ammonium glufosinate: resistant crops and resistance of weeds
Brazilian Journal of Herbicides, Brunharo, Caio, Christoffoleti, Pedro, Nicolai, Marcelo, 2014