Visiting Scientists, Borlaug Fellows, Fullbright Scholars
First name Last Name Year(s) Designator Home Country (from) Research
Leidy Paola Delgadillo Duran 2020 Borlaug Fellow Colombia Cd/ CAS9
Marieke Hilgers 2019-2020 Visiting Scientist Netherlands Functional genomics
Thierry Legrave 2019 Visiting Scientist France CRISPR disease genes
Gabriela Silva-Pumarada 2018 Visiting Scientist Puerto Rico Cacao
Fernando Santiago 2018 Visiting Scientist Puerto Rico Cacao
Adriana Gallego 2016-2017 Visiting Scientist Colombia Polyphenol light effect
Desire Pokou 2013, 2016, 2017 Visiting Scientist Ivory Coast Disease resistance in cacao
Annelle Holder-John 2016 Visiting Scientist Trinidad & Tobago Cutting evaluation in two species of cacao
Jaime Osorio 2016 Visiting Scientist Colombia Cadium uptake in cacao
Suzana Sato Guima 2015 Visiting Scientist Brazil Knowledge sharing
Lin Zhou 2015 Visiting Scientist China Defense gene in diversity population
Adrielle de Vasconcelos 2015 Visiting Scientist Brazil Knowledge sharing
Dadzie Abu 2013 Visiting Scientist Ghana Disease resistance in cacao
Luis Mejia-Franco 2009, 2012 Visiting Scientist Panama Cacao Endophytes
Claudia Arevalo 2012 Visiting Scientist Venezuela Cacao Transformation
Monica Santos-Mendoza 2011 Borlaug Fellow Mexico Transcriptomic and Motobolomic of Cocoa embriogenesis
Nicolas Dr. Niemenak 2010-2011 Fulbright Scholar Cameroon Maturation and germination physiology of cacao
Yaleidis Mendez 2010 Visiting Scientist Puerto Rico Cacao somatic embryogenesis
Rosalia Velasquez-Salazar 2008-2009 Visiting Scientist Venezuela Cacao and Jatropha Somatic Embryogenesis
Anna Muyiwa 2008 Visiting Scientist Nigeria Cacao, germplasm & germination
Antonio Figueira 2007-2008 Fulbright Scholar Brazil Global Analysis of Genes Differentially Expressed in the Cacao Genotypes in Response to Witches Broom Infection
Peter Aikpokpodion 2006-2007 Borlaug Fellow Nigeria Genetic Diversity in Cacao Collections in Nigeria Determined by Phenotypic and Simple Sequence Repeat Markers
Chia-Lock Tan 2006-2007 Fulbright Scholar Malasia Molecular and Biotechnological Techniques for the Improvement of Cocoa Against Pests
Catalina Chapparo 2001 Visiting Scientist Colombia Cacao Somatic Embryogenesis for Colombian Varieties

Post Doctoral Students
First name Last Name Year(s) Title Research PhD Institution
Jigeesha Mukhopadhyay 2024-2025 Research Assistant Cacao Ph.D. University of Manitoba, Canada
Patrick Thomas 2023-2025 Research Assistant Cacao Ph.D. University of California, Riverside
Sarah Prewitt 2017-2020 Research Assistant Cacao Ph. D. University of North Texas
Akiva Shalit-Kaneh 2017-2019 Research Assistant Cacao Ph.D. University of California, Davis
Andrew Fister 2016-2018 Research Assistant Cacao Ph.D. Penn State
Yufan Zhang 2015-2016 Research Assistant Cacao Ph.D. Penn State
Emily Helliwell 2013-2014 Research Assistant Cacao Ph.D. Penn State
Xiangling Shen 2011-2012 Research Assistant Cocoa propagation/ microarray analysis and bioinformatics Ph.D. Nanjing Agricultural University, China
Zi Shi 2010-2012 Research Assistant Bread Project/ Pathogense Ph.D. Penn State
Luis Mejia-Franco 2009-2010 Research Assistant Tree Symbiont Interactions Ph.D. Rutgers University
Marna Yandeau-Nelson 2005-2008 Research Assistant Maize Ph.D. Iowa State University
Joe Verica 2002-2007 Research Assistant Cacao Ph.D. Penn State
Jihong Li 2004-2005 Research Assistant Maize Ph.D. Hainan University, China
Yuan Yao 2002 Research Assistant Maize Ph.D. Jiangnan University, China
Susan Blauth 1997-1999 Research Assistant Maize Ph.D. Cornell University
Zhijian Li 1996-1998 Research Assistant Cacao Ph.D. Louisiana State University
Leslie Carlini 1994-1997 Research Assistant Plant Physiology Ph.D. Penn State
Siela Maximova 1996 Research Assistant Cacao Ph.D. Penn State
Lauren Gross 1992-1993 Research Assistant Plant Physiology Ph.D. Penn State
Nancy Eckardt 1992-1993 Research Assistant Maize Ph.D. Penn State

Graduate Students

Jishnu    Bhatt    2025
Ph.D. of Philosophy    Penn State    
Thesis: Enhancing the Efficiency of Somatic Embryogenesis and Genetic Transformation in Theobroma cacao L.

Francisco Miguel    Menéndez Burns    2025
Ph.D. Plant Biology    Penn State    
Thesis: Genomics and Physiological Response to Cadmium and the Evolution of Heavy Metal ATPases in Theobroma cacao L. 

Zach    Dashner    2024
Ph.D.  Horticulture    Penn State    
Thesis: Novel Biotechnological Approaches to Understand And Reduce Cadmium in the Iron Uptake Pathway of Theobroma Cacao

Daniel    Guarín    2024  Penn State 
Ph.D. dual-title in Soil Science and International Agriculture and Development       
Thesis: The Interplay of Bedrock Geomorphology and Human Impact on Soil Health: A Multiscale Assessment

Jhony     Benavides    2023  Penn State    
Ph.D. dual-title in Soil Science and International Agriculture and Development   
Thesis: Unmanned Aerial Vehicle Photogrammetry for Mapping Hydrologically Sensitive and Phosphorus Critical Source Areas

Winters    Noah    2022    Penn State
Ph.D. Ecology   
Thesis: Evolutionary and Functional Genetics of Disease Resistance in Tehobroma cacao and its Wild Relatives

Alejandro     Gil Aguirre    2022    Penn State
Ph.D. Education, Development, and Community Engagement
Thesis: Technology adoption by cacao farmers in San Vicente de Chucuri, Santander: the role of Cadmium regulations

Lina    Tami- Barrera    2021   Penn State
M.S. Dual title in Rural Sociology and International Agriculture & Development
Thesis: Socioeconomic and Technological Factors Influencing Technology Adoption in Cacao Farms of Two Post-conflict Regions in Colombia

Johann    Restrepo Rubio    2020    Penn State
M.S. Plant Sciences
Thesis: Characterization of the Function of Selected Phytophthora Megakarya Effector Proteins During Infection of Theobroma Cacao

Knollenberg    Benjamin    2020    Penn State
Ph.D. Plant Biology
Thesis: Characterization of Specialized Metabolites Involved in Black Pod Rot Resistance in Theobroma Cacao

Fister    Drew    2016    Penn State
Ph.D. in Genetics
Thesis: Functional Genomics of the Theobroma Cacao Defense Response

Sergio    Florez    2015    Penn State
Ph.D. in Chemical Engineering
Thesis: Improving Plant Propogation by Manipulating the Genetic and Physical Environment in Tissue Culture

Yufan    Zhang    2014     Penn State
Ph.D. in Plant Biology
Thesis: Functional genomics of Theobroma cacao fatty acid biosynthesis: Convergence of fatty acid desaturation, embryo development, and defense signaling responses

Zi     Shi    2010     Penn State
Ph.D. in Plant Biology
Thesis: Functional analysis of Non Expressor of pr1 (NPR1) and its paralog NPR3 in Theobroma cacao and Arabidopsis thaliana

Rachel     Melnick    2010    Penn State
Ph.D. in Plant Pathology
Thesis: Endophytic Bacillus SPP of Theobroma Cacao: Ecology and Potential for Biological Control of Cacao Diseases

Yi    Liu    2010    Penn State
Ph.D. in Integrated Biological Sciences
Thesis: Molecular Analysis of Genes Involved in the Synthesis of Proanthocyanidins in Theobroma in Cacao

Carter    Miller    2009    Penn State
Ph.D. in Integrated Biological Sciences
Thesis: An Integrated In Vitro and Greenhouse Orthotropic Clonal Propagation System for Theobroma Cacao L.

Huan    Xia    2009    Penn State
Ph.D. in Integrated Biological Sciences
Thesis: Structure and Function of Endosperm Starch from Maize Mutants Deficient in One or More Starch-branching Enzyme Isoform Activities

J.P.    Marelli    2008    Penn State
Ph.D. in Integrated Biological Sciences
Thesis: Solanum lycopersicum as a model system to study pathogenicity mechanisms of Moniliophthora perniciosa, the causal agent of witches' broom disease of Theobroma cacao

J.D.     Swanson    2005     Penn State
Ph.D. in Integrated Biological Sciences
Thesis: Flower development in Theobroma cacao L.: An assessment of morphological and molecular conservation of floral development between Arabidopsis thaliana and Theobroma cacao

Gabriela    Antunez De Mayolo    2003    Penn State
Ph.D. in Integrated Life Sciences
Thesis: Genetic engineering of Theobroma cacao and molecular studies on cacao defense responses

Melis    Cakirer    2003    Penn State
B.S. and M.S. in Food Sciences
Thesis: Color as an indicator of flavanol content in the fresh seeds of Theobroma cacao L

Jennifer    Tomscha    2001   Penn State
Ph.D. Horticulture
Thesis: Phosphatase Secretion Mutants in Arabidopsis Thaliana

Jennifer    Umpress    2001   Penn State
Ph.D. in Plant Physiology
Thesis: Phosphorus stress response in Arabidopsis

Abdoulaye    Traore    1999    Penn State
Ph.D. in Food Science
Thesis: Tissue culture and Transformation of cacao

Kyung-Nam    Kim    1997    Penn State
Ph.D. in Plant Physiology
Thesis: Isolation and promoter analysis of the genes for starch branching enzyme in maize

Ming    Gao    1996    Penn State
Ph.D. in Plant Physiology
Thesis: Molecular characterization of starch branching enzyme genes, SbeI, Sbe2b and Sbe2a in maize (Zea mays L.)

Siela     Maximova    1996    Penn State
Ph.D. in Plant Physiology
Thesis: Development of a genetic transformation system for apple

Dane    Fisher    1995    Penn State
Ph.D. in Genetics
Thesis: Molecular genetic analysis of multiple isoforms of starch branching enzyme with emphasis on Zea mays L.

Xiping    Niu    1995    Penn State
Ph.D. in Plant Physiology
Thesis: DNA binding specificity and interactions with nucleosomal DNA of the plant bZip protein EmBP-1

Sashi    Prabhakar    1995    Penn State
M.S. Plant Physiology
Thesis: cDNA cloning and characterization of a EmBP-1 homologue in maize (Zea mays): A basic leucine zipper transcription factor

Charlene    Chang    1993    Penn State
M.S. Plant Physiology
Thesis: Optimization of in vitro regeneration of geranium (Pelargonium xhortorum L.H. Bailey) for development of a genetic transformation system.

Mark    Guiltinan    1986 University of California, Irvine
Ph.D. Biology       
Thesis: The Isolation, Characterization and Intergeneric Transfer of Two Soybean (Glycine Max L.) Beta-tubulin Genes (TI Plasmid)

Staff
First name Last Name Year(s) Designator Title
Andrea Murphy-Faust 2017-2018 Staff Research Assistant
Teresa Soler 2014-2015 Staff Greenhouse/ lab Assistant
Matthew Dommel 2012-2013 Staff Research Assistant
Adam Clemens 2010-2013 Staff Research Assistant
Marissa Kenzakoski 2010-2013 Staff Research Assistant
Sharon Pishak 1996-2013 Staff Research Assistant, Lab Manager
Nur Sabrina Wahid 2010 Staff Research Assistant
Ann Young 1999-2010 Staff Research Assistant, Lab Manager
Melanie Schupp 2004-2005 Staff Research Assistant
Nianying Wang 2002-2003 Staff Research Assistant
Lisa Renshaw-Gegg 2001 Staff Research Assistant, Lab Manager
David Hulce 2000 Staff Research Assistant
Linda Miller 1999-2000 Staff Research Assistant, Lab Manager